Biomimetic flight and propulsion mechanisms Peer reviewed

Leaping out of the water: Aerial-aquatic locomotion with flapping wings

Raphael Zufferey, Simon Jeger, Moritz Hüsser, Fernando Ruíz and 3 more

Science | Jul 9, 2026

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It is shown that frequency adaptation, flexible wings, and powerful actuation enable seamless transitions without folding wings or legs, that large wings enhance flight without substantially reducing underwater efficiency, and that tail-body distance and egress angle affect water exit.

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Wing-propelled diving birds flap their wings to move through air and water, yet the wing morphology and kinematics that enable this behavior remain poorly understood because of the difficulty of collecting in situ data. The impact of flapping frequency, wing size, and stiffness on locomotion in—and transition between—the two media are still unknown. We compared data from diving birds against experiments using a flapping-wing robot capable of flying, swimming, plunge diving, and exiting the water. We show that frequency adaptation, flexible wings, and powerful actuation enable seamless transitions without folding wings or legs, that large wings enhance flight without substantially reducing underwater efficiency, and that tail-body distance and egress angle affect water exit. These results clarify how birds (and robots) balance multifluid locomotion constraints.

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Authors

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Raphael Zufferey

first | École Polytechnique Fédérale de Lausanne | ORCID 0000-0002-8722-6664

Simon Jeger

middle | École Polytechnique Fédérale de Lausanne | ORCID 0000-0003-3824-5612

Moritz Hüsser

middle | Massachusetts Institute of Technology

Fernando Ruíz

middle | École Polytechnique Fédérale de Lausanne | ORCID 0000-0003-0314-5478

Anthony B. Lapsansky

middle | Northwest Indian College | ORCID 0000-0001-7530-7830

Auke Jan Ijspeert

middle | École Polytechnique Fédérale de Lausanne | ORCID 0000-0003-1417-9980

Dario Floreano

last | École Polytechnique Fédérale de Lausanne | ORCID 0000-0002-5330-4863

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BibTeX

@article{Zufferey2026Leaping,
  title = {Leaping out of the water: Aerial-aquatic locomotion with flapping wings},
  author = {Raphael Zufferey and Simon Jeger and Moritz Hüsser and Fernando Ruíz and Anthony B. Lapsansky and Auke Jan Ijspeert and Dario Floreano},
  journal = {Science},
  year = {2026},
  doi = {10.1126/science.aeb6744},
  url = {https://doi.org/10.1126/science.aeb6744}
}

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